The Origin of Species - Charles Darwin (ebook audio reader txt) 📗

that earth occasionally adheres in some quantity to the feet and beaks of birds. Wading birds, which frequent the muddy edges of ponds, if suddenly flushed, would be the most likely to have muddy feet. Birds of this order wander more than those of any other; and are occasionally found on the most remote and barren islands of the open ocean; they would not be likely to alight on the surface of the sea, so that any dirt on their feet would not be washed off; and when gaining the land, they would be sure to fly to their natural freshwater haunts. I do not believe that botanists are aware how charged the mud of ponds is with seeds: I have tried several little experiments, but will here give only the most striking case: I took in February three tablespoonfuls of mud from three different points, beneath water, on the edge of a little pond; this mud when dry weighed only 6 and ¾ ounces; I kept it covered up in my study for six months, pulling up and counting each plant as it grew; the plants were of many kinds, and were altogether 537 in number; and yet the viscid mud was all contained in a breakfast cup! Considering these facts, I think it would be an inexplicable circumstance if waterbirds did not transport the seeds of freshwater plants to unstocked ponds and streams, situated at very distant points. The same agency may have come into play with the eggs of some of the smaller freshwater animals.

Other and unknown agencies probably have also played a part. I have stated that freshwater fish eat some kinds of seeds, though they reject many other kinds after having swallowed them; even small fish swallow seeds of moderate size, as of the yellow water-lily and Potamogeton. Herons and other birds, century after century, have gone on daily devouring fish; they then take flight and go to other waters, or are blown across the sea; and we have seen that seeds retain their power of germination, when rejected many hours afterwards in pellets or in the excrement. When I saw the great size of the seeds of that fine water-lily, the Nelumbium, and remembered Alph. de Candolle’s remarks on the distribution of this plant, I thought that the means of its dispersal must remain inexplicable; but Audubon states that he found the seeds of the great southern water-lily (probably according to Dr. Hooker, the Nelumbium luteum) in a heron’s stomach. Now this bird must often have flown with its stomach thus well stocked to distant ponds, and, then getting a hearty meal of fish, analogy makes me believe that it would have rejected the seeds in the pellet in a fit state for germination.

In considering these several means of distribution, it should be remembered that when a pond or stream is first formed, for instance on a rising islet, it will be unoccupied; and a single seed or egg will have a good chance of succeeding. Although there will always be a struggle for life between the inhabitants of the same pond, however few in kind, yet as the number even in a well-stocked pond is small in comparison with the number of species inhabiting an equal area of land, the competition between them will probably be less severe than between terrestrial species; consequently an intruder from the waters of a foreign country would have a better chance of seizing on a new place, than in the case of terrestrial colonists. We should also remember that many freshwater productions are low in the scale of nature, and we have reason to believe that such beings become modified more slowly than the high; and this will give time for the migration of aquatic species. We should not forget the probability of many freshwater forms having formerly ranged continuously over immense areas, and then having become extinct at intermediate points. But the wide distribution of freshwater plants, and of the lower animals, whether retaining the same identical form, or in some degree modified, apparently depends in main part on the wide dispersal of their seeds and eggs by animals, more especially by freshwater birds, which have great powers of flight, and naturally travel from one piece of water to another.

On the Inhabitants of Oceanic Islands

We now come to the last of the three classes of facts, which I have selected as presenting the greatest amount of difficulty with respect to distribution, on the view that not only all the individuals of the same species have migrated from some one area, but that allied species, although now inhabiting the most distant points, have proceeded from a single area, the birthplace of their early progenitors. I have already given my reasons for disbelieving in continental extensions within the period of existing species on so enormous a scale that all the many islands of the several oceans were thus stocked with their present terrestrial inhabitants. This view removes many difficulties, but it does not accord with all the facts in regard to the productions of islands. In the following remarks I shall not confine myself to the mere question of dispersal, but shall consider some other cases bearing on the truth of the two theories of independent creation and of descent with modification.

The species of all kinds which inhabit oceanic islands are few in number compared with those on equal continental areas: Alph. de Candolle admits this for plants, and Wollaston for insects. New Zealand, for instance, with its lofty mountains and diversified stations, extending over 780 miles of latitude, together with the outlying islands of Auckland, Campbell and Chatham, contain altogether only 960 kinds of flowering plants; if we compare this moderate number with the species which swarm over equal areas in Southwestern Australia or at the Cape of Good Hope, we must admit that some cause, independently of different physical conditions, has given rise to so great a difference in number. Even