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active in hard muscular work or hard thinking, while he feels passive in reflex action, in sensation, and in simply "being reminded" of anything without any effort on his own part. But he is active in everything he does, and he does everything that depends on his being alive. Life is activity, and every manifestation of life, such as reflex action or sensation, is a form of vital activity. The only way to be inactive is to be dead.

But vital activity is not "self-activity" in any absolute sense, for it is aroused by some stimulus. It does not issue from the individual as an isolated unit, but is his response to a stimulus. That is the sense of calling any mental process a reaction; it is something the individual does in response to a stimulus.

To call a sensation a form of reaction means, then, that the sensation is not something done to the person, nor passively received by him from outside, but something that he himself does when aroused to this particular form of activity. What comes from outside and is received by the individual is the stimulus, and the sensation is what he does in response to the stimulus. It represents the discharge of internal stored energy in a direction determined by his own inner mechanism. The sensation depends on his own make-up as well as on the nature of the stimulus, as is especially obvious when the sensation is abnormal or peculiar. Take the case of color blindness. The same stimulus that arouses in most people the sensation of red arouses in the color-blind individual the sensation of brown. Now what the color-blind individual receives, the light stimulus, is the same as what others receive, but he responds differently, i.e., with a different sensation, because his own sensory apparatus is peculiar.

The main point of this discussion is that all mental {47} phenomena, whether movements, sensations, emotions, impulses or thoughts, are a person's acts, but that every act is a response to some present stimulus. This rather obvious truth has not always seemed obvious. Some theorists, in emphasizing the spontaneity and "self-activity" of the individual, have pushed the stimulus away into the background; while others, fixing their attention on the stimulus, have treated the individual as the passive recipient of sensation and "experience" generally. Experience, however, is not received; it is lived, and that means done; only, it is done in response to stimuli. The concept of reaction covers the ground.

While speaking of sensations and thoughts as belonging under the general head of reactions, it is well, however, to bear in mind that all mental action tends to arouse and terminate in muscular and glandular activity. A thought or a feeling tends to "express itself" in words or (other) deeds. The motor response may be delayed, or inhibited altogether, but the tendency is always in that direction.

Different Sorts of Stimuli

To call all mental processes reactions means that it is always in order to ask for the stimulus. Typically, the stimulus is an external force or motion, such as light or sound, striking on a sense organ. There are also the internal stimuli, consisting of changes occurring within the body and acting on the sensory nerves that are distributed to the muscles, bones, lungs, stomach and most of the organs. The sensations of muscular strain and fatigue, and of hunger and thirst, are aroused by internal stimuli, and many reflexes are aroused in the same way.

Such internal stimuli as these are like the better known external stimuli in that they act upon sense organs; but it {48} seems necessary to recognize another sort of stimuli which act directly on the nerve centers in the brain. These may be called "central stimuli" and so contrasted with the "peripheral stimuli" that act on any sense organ, external or internal. To do this is to take considerable liberty with the plain meaning of "stimulus", and calls for justification. What is the excuse for thus expanding the notion of a stimulus?

The excuse is found in the frequent occurrence of mental processes that are not directly aroused by any peripheral stimulus, though they are plainly aroused by something else. Anything that arouses a thought or feeling can properly be called its stimulus. Now it often happens that a thought is aroused by another, just preceding thought; and it seems quite in order to call the first thought the stimulus and the second the response. A thought may arouse an emotion, as when the thought of my enemy, suddenly occurring to mind, makes me angry; the thought is then the stimulus arousing this emotional response.

If hearing you speak of Calcutta makes me think of India, your words are the stimulus and my thought the response. Well, then, if I think of Calcutta in the course of a train of thought, and next think of India, what else can we say than that the thought of Calcutta acts as a stimulus to arouse the thought of India as the response? In a long train of thought, where A reminds you of B and B of C and C of D, each of these items is, first, a response to the preceding, and, second, a stimulus to the one following.

There is no special difficulty with the notion of "central stimuli" from the physiological side. We have simply to think of one nerve center arousing another by means of the tract of axons connecting the two. Say the auditory center is aroused by hearing some one mention your friend's name, {49} and this promptly calls up a mental picture of your friend; here the auditory center has aroused the visual. What happens in a train of thought is that first one group of neurones is aroused to activity, and then this activity, spreading along the axons that extend from this group of neurones to another, arouses the second group to activity; and so on. The brain process may often be exceedingly complex, but this simple scheme gives the gist of it.

The way nerve currents must go shooting around the brain from one center or group of neurones to another, keeping it up for a long time without requiring any fresh peripheral stimulus, is remarkable. We have evidence of this sort of thing in a dream or fit of abstraction. Likely enough, the series of brain responses would peter out after awhile, in the absence of any fresh peripheral stimulus, and total inactivity ensue. But response of one brain center to nerve currents coming from another brain center, and not directly from any sense organ, must be the rule rather than the exception, since most of the brain neurones are not directly connected with any sense organ, but only with other parts of the brain itself. All the evidence we have would indicate that the brain is not "self-active", but only responsive; but, once thrown into activity at one point, it may successively become active at many other points, so that a long series of mental operations may follow upon a single sensory stimulus.

The Motor Centers, Lower and Higher

A "center" is a collection of nerve cells, located somewhere in the brain or cord, which gives off axons running to some other center or out to muscles or glands, while it also receives axons coming from other centers, or from sense organs. These incoming axons terminate in end-brushes and so form synapses with the dendrites of the local {50} nerve cells. The axons entering any center and terminating there arouse that center to activity, and this activity, when aroused, is transmitted out along the axons issuing from that center, and produces results where those axons terminate in their turn.



Fig. 12.--Side view of the left hemisphere of the brain, showing the motor and sensory areas (for the olfactory area, see Fig. 18). The visual area proper, or "visuo-sensory area," lies just around the corner from the spot marked "Visual," on the middle surface of the hemisphere, where it adjoins the other hemisphere. (Figure text: frontal lobe, parietal lobe, central fissure, occipital lobe, motor area, somesthetic area, auditory area, fissure of Sylvius, temporal lobe, brain stem, cerebellum)

The lower motor centers, called also reflex centers, are located in the cord or brain stem, and their nerve cells give rise to the axons that form the motor nerves and connect with the muscles and glands. A muscle is thrown into action by nerve currents from its lower motor center.

The principal higher motor center is the "motor area" of the brain, located in the cortex or external layer of gray matter, in the cerebrum. More precisely, the motor area is a long, narrow strip of cortex, lying just forward of what is called the "central fissure" or "fissure of Rolando".

{51}

If you run your finger over the top of the head from one side to the other, about halfway back from the forehead, the motor areas of the two cerebral hemispheres will lie close under the path traced by your finger.



Fig. 13.--(After Cajal.) Type of the brain cells that most directly control muscular movement. (Figure text: Axon. Giant pyramid cell from the motor area of the cerebral cortex, magnified 35 diameters. Cell body of same farther magnified)

The motor area in the right hemisphere is connected with the left half of the cord and so with the muscles of the left half of the body; the motor area of the left hemisphere similarly affects {52} the right half of the body. Within the motor area are centers for the several limbs and other motor organs. Thus, at the top, near the middle line of the head (and just about where the phrenologists located their "bump of veneration"!), is the center for the legs; next below and to the side is the center for the trunk, next that for the arm, next that for head movements, and at the bottom, not far from the ears, is the center for tongue and mouth.



Fig. 14.--The nerve path by which the motor area of the cortex influences the muscles. The upper part of this path, consisting of axons issuing from the giant pyramids of the motor area and extending down into the spinal cord, is the pyramidal tract. The lower part of the path consists of axons issuing from the motor cells of the cord and extending out to the muscles. The top of the figure represents a vertical cross-section of the brain, such as is given, on a larger scale, in Fig. 18. (Figure text: cortex, cord, muscles)

The largest nerve cells of all are found in the motor area, and are called, from their shape, the "giant pyramids". They have large dendrites and very long axons, which latter, {53} running in a thick bundle down from the cortex through the brain stem and cord, constitute the "pyramidal tract", the principal path of communication from the cerebrum to the lower centers. The motor area of the brain has no direct connection with any muscle, but acts through the pyramidal tract on the lower centers, which in turn act on the muscles.

How The Brain Produces Muscular Movements

The motor area is itself aroused to action by nerve currents entering it through axons coming from other parts of the cortex; and it is by way of the motor area that any other part of the cortex produces bodily movement. There are a few exceptions, as, for example, the movements of the eyes are produced generally by the "visual area" acting directly on the lower motor centers for the eye in the brain stem; but, in the main, any motor effect of brain action is exerted through the motor area. The motor area, as already mentioned, acts on the lower motor centers in the cord and brain stem, and these in turn on the muscles; but we must look into this matter a little more closely.

A lower motor center is a group of motor

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