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fishes (the teleosts) the tail fin is not vertebrated: the spinal column ends in a broad plate, to which the diverging fin rays are attached.

But along with these embryonic characters, which were common to all Devonian fishes, there were other structures in certain groups which foreshadowed the higher structures of the land vertebrates which were yet to come: air sacs which were to develop into lungs, and cartilaginous axes in the side fins which were a prophecy of limbs. The vertebrates had already advanced far enough to prove the superiority of their type of structure to all others. Their internal skeleton afforded the best attachment for muscles and enabled them to become the largest and most powerful creatures of the time. The central nervous system, with the predominance given to the ganglia at the fore end of the nerve cord,—the brain,— already endowed them with greater energy than the invertebrates; and, still more important, these structures contained the possibility of development into the more highly organized land vertebrates which were to rule the earth.

TELEOSTS. The great group of fishes called the teleosts, or those with complete bony skeletons, to which most modern fishes belong, may be mentioned here, although in the Devonian they had not yet appeared. The teleosts are a highly specialized type, adapted most perfectly to their aquatic environment. Heavy armor has been discarded, and reliance is placed instead on swiftness. The skeleton is completely ossified and the notochord removed. The vertebrae have been economically withdrawn from the tail, and the cartilaginous axis of the side fins has been fotfoid unnecessary. The air sac has become a swim bladder. In this complete specialization they have long since lost the possibility of evolving into higher types.

It is interesting to note that the modern teleosts in their embryological growth pass through the stages which characterized the maturity of their Devonian ancestors; their skeleton is cartilaginous and their tail fin vertebrated.

CHAPTER XIX THE CARBONIFEROUS

The Carboniferous system is so named from the large amount of coal which it contains. Other systems, from the Devonian on, are coal bearing also, but none so richly and to so wide an extent. Never before or since have the peculiar conditions been so favorable for the formation of extensive coal deposits.

With few exceptions the Carboniferous strata rest on those of the Devonian without any marked unconformity; the one period passed quietly into the other, with no great physical disturbances.

The Carboniferous includes three distinct series. The lower is called the MISSISSIPPIAN, from the outcrop of its formations along the Mississippi River in central and southern Illinois and the adjacent portions of Iowa and Missouri. The middle series is called the PENNSYLVANIAN (or Coal Measures), from its wide occurrence over Pennsylvania. The upper series is named the PERMIAN, from the province of Perm in Russia.

THE MISSISSIPPIAN SERIES. In the interior the Mississippian is composed chiefly of limestones, with some shales, which tell of a clear, warm, epicontinental sea swarming with crinoids, corals, and shells, and occasionally clouded with silt from the land.

In the eastern region, New York had been added by uplift to the Appalachian land which now was united to the northern area. From eastern Pennsylvania southward there were laid in a subsiding trough, first, thick sandstones (the Pocono sandstone), and later still heavier shales,—the two together reaching the thickness of four thousand feet and more. We infer a renewed uplift of Appalachia similar to that of the later epochs of the Devonian, but as much less in amount as the volume of sediments is smaller.

THE PENNSYLVANIAN SERIES

The Mississippian was brought to an end by a quiet oscillation which lifted large areas slightly above the sea, and the Pennsylvanian began with a movement in the opposite direction. The sea encroached on the new land, and spread far and wide a great basal conglomerate and coarse sandstones. On this ancient beach deposit a group of strata rests which we must now interpret. They consist of alternating shales and sandstones, with here and there a bed of limestone and an occasional seam of coal. A stratum of fire clay commonly underlies a coal seam, and there occur also beds of iron ore. We give a typical section of a very small portion of the series at a locality in Pennyslvania. Although some of the minor changes are omitted, the section shows the rapid alternation of the strata:

Feet 9 Sandstone and shale . . . . . . . . 25 8 Limestone . . . . . . . . . . . . . 18 7 Sandstone . . . . . . . . . . . . . 10 6 Coal . . . . . . . . . . . . . . . 1-6 5 Shale . . . . . . . . . . . . . . . 0-2 4 Sandstone . . . . . . . . . . . . . 40 3 Limestone . . . . . . . . . . . . . 10 2 Coal . . . . . . . . . . . . . . . 5-12 1 Fire clay . . . . . . . . . . . . . 3

This section shows more coal than is usual; on the whole, coal seams do not take up more than one foot in fifty of the Coal Measures. They vary also in thickness more than is seen in the section, some exceptional seams reaching the thickness of fifty feet.

HOW COAL WAS MADE.

1. Coal is of vegetable origin. Examined under the microscope even anthracite, or hard coal, is seen to contain carbonized vegetal tissues. There are also all gradations connecting the hardest anthracite—through semibituminous coal, bituminous or soft coal, lignite (an imperfect coal in which sometimes woody fibers may be seen little changed)—with peat and decaying vegetable tissues. Coal is compressed and mineralized vegetal matter. Its varieties depend on the perfection to which the peculiar change called bituminization has been carried, and also, as shown in the table below, on the degree to which the volatile substances and water have escaped, and on the per cent of carbon remaining.

                       Peat Lignite Bituminous Coal
Anthracite
                   Dismal Swamp Texas Penn.
Penn.
    Moisture . . . . 78.89 14.67 1.30 2.74
    Volatile matter . 13.84 37.32 20.87 4.25
    Fixed carbon . . 6.49 41.07 67.20 81.51
    Ash . . . . . . . 0.78 6.69 8.80 10.87

2. The vegetable remains associated with coal are those of land plants.

3. Coal accumulated in the presence of water; for it is only when thus protected from the air that vegetal matter is preserved.

4. The vegetation of coal accumulated for the most part where it grew; it was not generally drifted and deposited by waves and currents. Commonly the fire clay beneath the seam is penetrated with roots, and the shale above is packed with leaves of ferns and other plants as beautifully pressed as in a herbarium. There often is associated with the seam a fossil forest, with the stumps, which are still standing where they grew, their spreading roots, and the soil beneath, all changed to stone. In the Nova Scotia field, out of seventy-six distinct coal seams, twenty are underlain by old forest grounds.

The presence of fire clay beneath a seam points in the same direction. Such underclays withstand intense heat and are used in making fire brick, because their alkalies have been removed by the long-continued growth of vegetation.

Fuel coal is also too pure to have been accumulated by driftage. In that case we should expect to find it mixed with mud, while in fact it often contains no more ash than the vegetal matter would furnish from which it has been compressed.

These conditions are fairly met in the great swamps of river plains and deltas and of coastal plains, such as the great Dismal Swamp, where thousands of generations of forests with their undergrowths contribute their stems and leaves to form thick beds of peat. A coal seam is a fossil peat bed.

GEOGRAPHICAL CONDITIONS DURING THE PENNSYLVANIAN. The Carboniferous peat swamps were of vast extent. A map of the Coal Measures (Fig. 260) shows that the coal marshes stretched, with various interruptions of higher ground and straits of open water, from eastern Pennsylvania into Alabama, Texas, and Kansas. Some individual coal beds may still be traced over a thousand square miles, despite the erosion which they have suffered. It taxes the imagination to conceive that the varied region included within these limits was for hundreds of thousands of years a marshy plain covered with tropical jungles such as that pictured in Figure 304.

On the basis that peat loses four fifths of its bulk in changing to coal, we may reckon the thickness of these ancient peat beds. Coal seams six and ten feet thick, which are not uncommon, represent peat beds thirty and fifty feet in thickness, while mammoth coal seams fifty feet thick have been compressed from peat beds two hundred and fifty feet deep.

At the same time, the thousands of feet of marine and freshwater sediments, with their repeated alternations of limestones, sandstones, and shales, in which the seams of coal occur, prove a slow subsidence, with many changes in its rate, with halts when the land was at a stillstand, and with occasional movements upward.

When subsidence was most rapid and long continued the sea encroached far and wide upon the lowlands and covered the coal swamps with sands and muds and limy oozes. When subsidence slackened or ceased the land gained on the sea. Bays were barred, and lagoons as they gradually filled with mud became marshes. River deltas pushed forward, burying with their silts the sunken peat beds of earlier centuries, and at the surface emerged in broad, swampy flats,—like those of the deltas of the Mississippi and the Ganges,—which soon were covered with luxuriant forests. At times a gentle uplift brought to sea level great coastal plains, which for ages remained mantled with the jungle, their undeveloped drainage clogged with its debris, and were then again submerged.

PHYSICAL GEOGRAPHY OF THE SEVERAL REGIONS. THE ACADIAN REGION lay on the eastern side of the northern land, where now are New Brunswick and Nova Scotia, and was an immense river delta. Here river deposits rich in coal accumulated to a depth of sixteen thousand feet. The area of this coal field is estimated at about thirty-six thousand square miles.

THE APPALACHIAN REGION skirts the Appalachian oldland on the west from the southern boundary of New York to northern Alabama, extending west into eastern Ohio. The Cincinnati anticline was now a peninsula, and the broad gulf which had lain between it and Appalachia was transformed at the beginning of the Pennsylvanian into wide marshy plains, now sinking beneath the sea and now emerging from it. This area subsided during the Carboniferous period to a depth of nearly ten thousand feet.

THE CENTRAL REGION lay west of the peninsula of the Cincinnati anticline, and extended from Indiana west into eastern Nebraska, and from central Iowa and Illinois southward about the ancient island in Missouri and Arkansas into Oklahoma and Texas. On the north the subsidence in this area was comparatively slight, for the Carboniferous strata scarcely exceed two thousand feet in thickness. But in Arkansas and Indian Territory the downward movement amounted to four and five miles, as is proved by shoal water deposits of that immense thickness.

The coal fields of Indiana, and Illinois are now separated by erosion from those lying west of the Mississippi River. At

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